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Ganga-Brahmaputra-Meghna basin, India, Bhutan and Nepa
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Gin River basin, Sinharaja World Heritage Site, Sri Lanka
(Photo from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Diagnosis: Devario annnataliae is distinguished from all Sri Lankan Devario by the combination of following
characters: body depth 24–27 %SL; snout length equal to or greater than eye diameter; 5 or 6 irregular, dark
blue vertical bars on anterior half of body; P stripe, originating just anterior to origin of dorsal-fin; 16–17
predorsal scales; 10˝ branched dorsal-fin rays; 11˝–14˝ branched anal-fin rays; a minute process on first
infraorbital; danionin notch present; dorsal fin origin to hypural distance, when carried forward, falling well
short of posterior border of eye; pectoral-fin, when adpressed, just reaching pelvic-fin origin.
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Chittagong Division, Bangladesh.
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Chindwin River drainage, Manipur, India and Myanma
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Bangladesh, India, Myanmar, Nepal and Pakistan
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Border between Myanmar and China
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Kerala, India and Sri Lanka; introduced elsewhere
(Photo and diagnosis from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Diagnosis: Devario malabaricus is distinguished from all other members of the genus by the combination of the
following characters: no process on 1st infraorbital; body depth 27–35 %SL; predorsal scales15–17; branched
dorsal-fin rays11˝–12˝; branched anal-fin rays 12˝–17˝; danionin notch present; dorsal fin origin to hypural
distance when carried forward falling well short of posterior border of eye; pectoral-fin tip almost reaching
pelvic-fin origin when adpressed; snout length subequal to or greater than eye diameter; P stripe originating
level with origin of pelvic-fin, 1–2 scale-widths anterior to dorsal-fin origin; P-1 stripe less than half width of
P stripe, bifurcated anteriorly by a more or less broken whitish line; anterior half of body with 5–6 dark,
irregular, vertical bars. No nuptial tubercles in both sexes.
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2020 Sudasinghea, Pethiyagoda & Meegaskumbura
Sri Lanka, Sudasinghea, Pethiyagoda & Meegaskumbura
Devario memorialis Sudasinghea, Pethiyagoda & Meegaskumbura 2020 On 14 May 2016, a massive low-pressure system formed over the Bay of Bengal, causing torrential rainfall across Sri Lanka. It strengthened to form Cyclone Roanu. Over the next four days, the rain caused devastating floods and landslides across the eastern and southern parts of the country. According to Wikipedia, 101 people died and 100 are still missing. This species was named in honor of those killed during that Cyclone.
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Kelaniweli Forest Reserve, Sri Lanka
(Photo and diagnosis from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Diagnosis: Devario micronema is distinguished from all other species of the genus by the combination of the
following characters: body depth 29–30 %SL; snout length equal to or greater than eye diameter; 3–5 irregular,
vertical bars on anterior half of body; P stripe broader than P+1 and P-1 stripes; 14–17 predorsal scales; 10–11
branched dorsal-fin rays; 12˝–14˝ branched anal-fin rays; a prominent, square-shaped process on posterior
edge of first infraorbital; danionin notch present; dorsal fin origin to hypural distance, when carried forward,
reaching to reaching posterior border of eye; pectoral-fin, adpressed, just reaching pelvic-fin origin.
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Mahaweli River basin, central Sri Lanka
(Photo from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Given its preoccupation in Devario by Leuciscus lineolatus Blyth, 1858 (Fang Kullander 2001; Kottelat 2013), Batuwita et. al. in 2017 provided the replacement name Devario monticola, nomen novum, for Danio lineolatus Bleeker, 1863.
Diagnosis: Devario monticola is distinguished from all other species of Sri Lankan Devario by the combination
of following characters: body depth 26–29 %SL; snout length equal to or greater than eye diameter; 4 or 5 dark,
irregular, vertical bars restricted to anterior half of body; P-1 stripe narrower than P stripe, originating level with
P stripe, extending to origin of caudal peduncle; 13–19 predorsal scales; 9˝–11˝ branched dorsa-fin rays, 12˝–
15˝ branched anal-fin rays; a process on first infraorbital; danionin notch present; dorso-hypural distance when
carried forward falling well short of posterior border of eye; and pectoral-fin, when adpressed, not reaching
pelvic-fin origin.
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Nilwala basin, southwest Sri Lanka
(Photo from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Diagnosis: Devario pathirana is distinguished from all other species of the genus by the combination of the
following characters: 8–9 dark, irregular bars on body between pectoral-fin origin and caudal-peduncle origin;
P+1 and P-1 stripes absent; body depth 30–33 %SL; snout length less than eye diameter; 14–15 predorsal scales;
11˝–12˝ branched dorsal-fin rays; 14˝ branched anal-fin rays; a blunt, knob-like process on 1st infraorbital;
danionin notch present; dorsal fin origin to hypural distance when carried forward reaching middle of eye;
pectoral-fin, when adpressed, just reaching pelvic-fin origin.
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Thanlwin basin, Myanmar and China
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Gin River basin, Sri Lanka.
(Photo from Batuwita et. al. 2017 monograph on Sri Lankan Devario)
Diagnosis: Devario udenii is distinguished from all other species of the genus by the combination of following
characters: body depth 26–36 %SL; snout length less than eye diameter; colour pattern consisting of 5–6 dark,
irregular, broad, vertical bars anterior to level of dorsal-fin origin; 14–17 predorsal scales; 9˝–12˝ branched
dorsal-fin rays and 12˝–15˝ branched anal-fin rays; danionin notch present; dorsal fin origin to hypural
distance, when carried forward, reaching middle of eye; pectoral-fin, when adpressed, reaching pelvic-fin origin;
P-1 stripe subequal in width to P stripe.
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Border areas of Manipur, India and Myanmar
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